phd biochemistry courses

• Biochemistry and Molecular Biology
• Biostatistics
• Environmental Health and Engineering
• Epidemiology
• Health Policy and Management
• Health, Behavior and Society
• International Health
• Mental Health
• Molecular Microbiology and Immunology
• Population, Family and Reproductive Health
• Program Finder
• Admissions Services
• Course Directory
• Academic Calendar
• Hybrid Campus
• Lecture Series
• Convocation
• Strategy and Development
• Implementation and Impact
• Integrity and Oversight
• In the School
• In the Field
• In Baltimore
• Resources for Practitioners
• Articles & News Releases
• In The News
• Statements & Announcements
• At a Glance
• Student Life
• Strategic Priorities
• Inclusion, Diversity, Anti-Racism, and Equity (IDARE)
• What is Public Health?

Doctor of Philosophy (PhD)

Offered By: Department of Biochemistry and Molecular Biology

Onsite | Full-Time | 5 – 6 years

• MSPH Field Placements
• Master's Essays
• MAS Application Fee Waiver Requirements
• Master of Arts and Master of Science in Public Health (MA/MSPH)
• Master of Arts in Public Health Biology (MAPHB)
• Master of Bioethics (MBE)
• Mission, Vision, and Values
• Student Experience
• Program Outcomes
• For Hopkins Undergraduate Students
• Master of Health Science (MHS) - Department of Biochemistry and Molecular Biology
• Master of Health Science (MHS) - Department of Epidemiology
• Alumni Update
• MHS Combined with a Certificate Program
• Master of Health Science (MHS) - Department of Molecular Microbiology and Immunology
• Bachelor's/MHS in Health Economics and Outcomes Research
• MHS HEOR Careers
• Frequently Asked Questions
• Master of Health Science (MHS)
• Concurrent School-Wide Master of Health Science Program in Biostatistics
• Master of Health Science - Department of Population, Family and Reproductive Health
• Master of Health Science Online (MHS) - Department of Population, Family and Reproductive Health
• Careers in Health Economics
• Core Competencies
• Meet the Director
• What is Health Economics
• MPH Capstone Schedule
• Concentrations
• Online/Part-Time Format
• Requirements

Tuition and Funding

• Executive Board Faculty
• Master of Science (ScM) - Department of Biochemistry and Molecular Biology
• Master of Science (ScM) - Department of Biostatistics
• Master of Science (ScM) - Department of Epidemiology
• Master of Science (ScM) - Department of Molecular Microbiology and Immunology
• Bachelor's/MSPH in Health Policy
• FAQ for MSPH in Health Policy
• Field Placement Experience
• MSPH Capstone
• MSPH Practicum
• Required and Elective Courses
• Student Timeline
• Career Opportunities
• 38-Week Dietetics Practicum
• Completion Requirements
• MSPH/RD Program FAQ
• Program Goals
• Application Fee Waiver Requirements
• Doctor of Philosophy (PhD) - Department of Biostatistics
• Doctor of Philosophy (PhD) - Department of Epidemiology
• Program Goals and Expectations
• Doctor of Philosophy (PhD) - Department of Molecular Microbiology and Immunology
• Doctor of Philosophy (PhD) - Department of Population, Family and Reproductive Health
• Doctor of Philosophy (PhD) in Clinical Investigation
• Recent Graduates and Dissertation Titles
• PhD Funding
• PhD TA Requirement
• Recent Dissertation Titles
• JHU-Tsinghua Doctor of Public Health
• Prerequisites
• Concentration in Women’s and Reproductive Health
• Custom Track
• Concentration in Environmental Health
• Concentration in Global Health: Policy and Evaluation
• Concentration in Health Equity and Social Justice
• Concentration in Health Policy and Management
• Concentration in Implementation Science
• Combined Bachelor's / Master's Programs
• Concurrent MHS Option for BSPH Doctoral Students
• Concurrent MSPH Option for JHSPH Doctoral students
• Doctor of Medicine and Doctor of Philosophy (MD/PhD)
• Adolescent Health Certificate Program
• Bioethics Certificate Program
• Clinical Trials Certificate Program
• Community- Based Public Health Certificate Program
• Demographic Methods Certificate Program
• Epidemiology for Public Health Professionals Certificate Program
• Evaluation: International Health Programs Certificate Program
• Frequently Asked Questions for Certificate Programs
• Gender and Health Certificate Program
• Gerontology Certificate Program
• Global Digital Health Certificate Program
• Global Health Certificate Program
• Global Health Practice Certificate Program
• Health Communication Certificate Program
• Health Disparities and Health Inequality Certificate Program
• Health Education Certificate Program
• Health Finance and Management Certificate Program
• Health and Human Rights Certificate Program
• Healthcare Epidemiology and Infection Prevention and Control Certificate Program
• Humanitarian Health Certificate Program
• Implementation Science and Research Practice Certificate Program
• Injury and Violence Prevention Certificate Program
• International Healthcare Management and Leadership Certificate Program
• Leadership for Public Health and Healthcare Certificate Program
• Lesbian, Gay, Bisexual, Transgender, and Queer (LGBTQ) Public Health Certificate Program
• Maternal and Child Health Certificate Program
• Mental Health Policy, Economics and Services Certificate Program
• Non-Degree Students General Admissions Info
• Pharmacoepidemiology and Drug Safety Certificate Program
• Population Health Management Certificate Program
• Population and Health Certificate Program
• Public Health Advocacy Certificate Program
• Public Health Economics Certificate Program
• Public Health Informatics Certificate Program
• Public Health Practice Certificate Program
• Public Health Training Certificate for American Indian Health Professionals
• Public Mental Health Research Certificate Program
• Quality, Patient Safety and Outcomes Research Certificate Program
• Quantitative Methods in Public Health Certificate Program
• Requirements for Successful Completion of a Certificate Program
• Rigor, Reproducibility, and Responsibility in Scientific Practice Certificate Program
• Risk Sciences and Public Policy Certificate Program
• Spatial Analysis for Public Health Certificate Program
• Training Certificate in Public Health
• Tropical Medicine Certificate Program
• Tuition for Certificate Programs
• Vaccine Science and Policy Certificate Program
• Online Student Experience
• MAS and Affiliated Certificate Programs
• Barcelona Information
• Registration, Tuition, and Fees
• Agency Scholarship Application
• General Scholarship Application
• UPF Scholarship Application
• Course Evaluations
• Online Courses
• Registration
• General Institute Tuition Information
• International Students
• Directions to the Bloomberg School
• All Courses
• Important Guidance for ONSITE Students
• D.C. Courses
• Registration and Fees
• Cancellation and Closure Policies
• Application Procedures
• Career Search
• Current Activities
• Current Trainees
• Related Links
• Process for Appointing Postdoctoral Fellows
• Message from the Director
• Program Details
• Admissions FAQ
• Current Residents
• Elective Opportunities for Visiting Trainees
• What is Occupational and Environmental Medicine?
• Admissions Info
• Graduates by Year
• Compensation and Benefits
• How to Apply
• Academic Committee
• Course Details and Registration
• Tuition and Fees
• ONLINE SOCI PROGRAM
• Principal Faculty
• General Application
• JHHS Application
• Our Faculty
• Descripción los Cursos
• Programa en Epidemiología para Gestores de Salud, Basado en Internet
• Consultants
• Britt Dahlberg, PhD
• Joke Bradt, PhD, MT-BC
• Mark R. Luborsky, PhD
• Marsha Wittink, PhD
• Rebekka Lee, ScD
• Su Yeon Lee-Tauler, PhD
• Theresa Hoeft, PhD
• Vicki L. Plano Clark, PhD
• Program Retreat
• Mixed Methods Applications: Illustrations
• Announcements
• 2023 Call for Applications
• Jennifer I Manuel, PhD, MSW
• Joke Bradt, PhD
• Josiemer Mattei, PhD, MPH
• Justin Sanders, MD, MSc
• Linda Charmaran, PhD
• Nao Hagiwara, PhD
• Nynikka R. A. Palmer, DrPH, MPH
• Olayinka O. Shiyanbola, BPharm, PhD
• Sarah Ronis, MD, MPH
• Susan D. Brown, PhD
• Tara Lagu, MD, MPH
• Theresa Hoft, PhD
• Wynne E. Norton, PhD
• Yvonne Mensa-Wilmot, PhD, MPH
• A. Susana Ramírez, PhD, MPH
• Animesh Sabnis, MD, MSHS
• Autumn Kieber-Emmons, MD, MPH
• Benjamin Han, MD, MPH
• Brooke A. Levandowski, PhD, MPA
• Camille R. Quinn, PhD, AM, LCSW
• Justine Wu, MD, MPH
• Kelly Aschbrenner, PhD
• Kim N. Danforth, ScD, MPH
• Loreto Leiva, PhD
• Marie Brault, PhD
• Mary E. Cooley, PhD, RN, FAAN
• Meganne K. Masko, PhD, MT-BC/L
• PhuongThao D. Le, PhD, MPH
• Rebecca Lobb, ScD, MPH
• Allegra R. Gordon, ScD MPH
• Anita Misra-Hebert, MD MPH FACP
• Arden M. Morris, MD, MPH
• Caroline Silva, PhD
• Danielle Davidov, PhD
• Hans Oh, PhD
• J. Nicholas Dionne-Odom, PhD RN ACHPN
• Jacqueline Mogle, PhD
• Jammie Hopkins, DrPH, MS
• Joe Glass, PhD MSW
• Karen Whiteman, PhD MSW
• Katie Schultz, PhD MSW
• Rose Molina, MD
• Uriyoán Colón-Ramos, ScD MPA
• Andrew Riley, PhD
• Byron J. Powell, PhD, LCSW
• Carrie Nieman MD, MPH
• Charles R. Rogers, PhD, MPH, MS, CHES®
• Emily E. Haroz, PhD
• Jennifer Tsui, Ph.D., M.P.H.
• Jessica Magidson, PhD
• Katherine Sanchez, PhD, LCSW
• Kelly Doran, MD, MHS
• Kiara Alvarez, PhD
• LaPrincess C. Brewer, MD, MPH
• Melissa Radey, PhD, MA, MSSW
• Sophia L. Johnson, PharmD, MPH, PhD
• Supriya Gupta Mohile, MD, MS
• Virginia McKay, PhD
• Andrew Cohen, MD, PhD
• Angela Chen, PhD, PMHNP-BC, RN
• Christopher Salas-Wright, PhD, MSW
• Eliza Park MD, MS
• Jaime M. Hughes, PhD, MPH, MSW
• Johanne Eliacin, PhD, HSPP
• Lingrui Liu ScD MS
• Meaghan Kennedy, MD
• Nicole Stadnick, PhD, MPH
• Paula Aristizabal, MD
• Radhika Sundararajan, MD
• Sara Mamo, AuD, PhD
• Tullika Garg, MD MPH FACS
• Allison Magnuson, DO
• Ariel Williamson PhD, DBSM
• Benita Bamgbade, PharmD, PhD
• Christopher Woodrell MD
• Hung-Jui (Ray) Tan, MD, MSHPM
• Jasmine Abrams, PhD
• Jose Alejandro Rauh-Hain, MD
• Karen Flórez, DrPH, MPH
• Lavanya Vasudevan, PhD, MPH, CPH
• Maria Garcia, MD, MPH
• Robert Brady, PhD
• Saria Hassan, MD
• Scherezade Mama, DrPH
• Yuan Lu, ScD
• 2021 Scholars
• Sign Up for Our Email List
• Workforce Training
• Cells-to-Society Courses
• Course/Section Numbers Explained
• Pathway Program with Goucher College
• The George G. Graham Lecture

About the PhD in Biochemistry and Molecular Biology Program

In the Biochemistry and Molecular Biology PhD program, faculty, and students work together to increase knowledge of the biochemical and molecular bases of normal and abnormal cellular processes. Our program trains students to be successful independent scientists and gives them the knowledge, research training, and leadership skills to continue to provide new insights into the biomedical issues that have a profound impact on public health. Cancer biology is a historical and continuing area of interest for many faculty in our program, which has been supported in part by a training grant from the National Institutes of Health’s National Cancer Institute since 1975.

Students engage in a rigorous course curriculum and a range of structured and informal activities outside the classroom and lab to build their skills. They will pursue their thesis research in the lab of one of our over forty training faculty across the Johns Hopkins Bloomberg School of Public Health and the Johns Hopkins School of Medicine.

Visit our dedicated PhD program website to learn more about the diverse research training opportunities of the program.

PhD in Biochemistry and Molecular Biology Program Highlights

Our position within the School of Public Health provides a unique setting in which students learn how biochemistry, molecular biology, physical chemistry, cell biology, and genetics can be used to solve significant problems in public health and medicine. Our program offers:

• Training faculty from across the School of Public Health and the School of Medicine
• A strong grounding in the science of biomedical and public health research through a core curriculum that includes courses taught by leading experts from the Schools of Public Health and Medicine
• Training outside the lab and classroom in key skills such as communications and leadership
• Opportunities to build strong communications skills through a range of speaking venues including journal club, research colloquium, department retreats, and national meetings
• Teaching Assistant service to build teaching and interpersonal skills, with options for additional training and professional development through the Johns Hopkins University Teaching Academy to further develop skills
• Access to the Johns Hopkins School of Medicine Professional Development and Career Office , offering excellent career services and professional development,  including the BMB-required OPTIONS program, a guided process of career exploration for paths from medicine to biotech to academia and beyond
• Opportunities to participate in community service and outreach, with a focus on our East Baltimore neighborhoods, through the Johns Hopkins University community engagement and service-learning center, SOURCE

Training faculty across the School of Public Health and the School of Medicine

Schools that students can take courses in: Public Health, Arts & Sciences, Medicine, and Engineering

Two-month rotations in the first year prior to selecting thesis lab

Average number of incoming students in the BMB PhD degree program each year

What Can You Do With a PhD In Biochemistry And Molecular Biology?

The Biochemistry and Molecular Biology PhD program prepares students for a range of biomedical and health sciences careers, including in academia, industry, policy, and beyond. Visit the Graduate Employment Outcomes Dashboard to learn about Bloomberg School graduates' employment status, sector, and salaries.

Sample Careers

• Research Scientist
• Science Policy Adviser
• Biotech Executive
• Senior Scientist
• Patent Lawyer
• Science Policy Analyst/Advocate
• Science Writer/Journalist
• Biological Sciences Teacher

Topic Areas

The BMB PhD program faculty conduct research to gain new insights into the cellular and molecular mechanisms underlying normal and abnormal cellular processes, and their relevance as targets for improving health and treating disease. Our training program places particular emphasis on mechanistic approaches to research problems, and cancer biology has had a prominent place in our research interest for over 50 years .

Common topic areas within our faculty's diverse research interests include:

• Biophysics and Structural Biology
• Cancer Biology
• Chemical Biology and Proteomics
• Cell Biology
• Cellular Stress and Cell Signaling
• Genetics, Genomics, and Gene Regulation
• Immunology and Infectious Diseases
• Translational Research

Curriculum for the PhD in Biochemistry and Molecular Biology

The BMB PhD offers students a rigorous course curriculum, including a set of common core classes from the Schools of Public Health and Medicine. Students further tailor their curriculum with elective courses chosen based on interests and career goals, with options that span the Schools of Public Health, Medicine, Engineering, and Arts and Sciences. A rich array of seminar programs and journal clubs are also available to all students.

Browse an overview of the requirements for this PhD program in the JHU  Academic Catalogue  and explore all course offerings in the Bloomberg School  Course Directory .

Admissions Requirements

For the general admissions requirements see our How to Apply page. The specific program also requires:

Prior Research Experience

Laboratory research experience (from academia, industry, etc.) is required

Prior Coursework

Strong background in the sciences, particularly in chemistry, biochemistry, or biology

Standardized Test Scores

Standardized test scores (GRE) are optional for this program. The admissions committee will make no assumptions if a standardized test score is omitted from an application, but will require evidence of quantitative/analytical ability through other application components such as academic transcripts and/or supplemental questions.  Applications will be reviewed holistically based on all application components.

Program Faculty Spotlight

Ashani T. Weeraratna

Ashi Weeraratna, PhD, studies how cancer cells move to distant sites and how changes in the normal cells around a tumor contribute to their movement, especially as we age.

Michael J. Matunis

Michael Matunis, PhD, studies how protein modification by SUMO—the small ubiquitin-related modifier—drives changes in key cellular pathways from stress response to DNA repair.

Jennifer M. Kavran

Jennifer Kavran, PhD, MS, MPhil, is a biophysicist who investigates how cells communicate with each other and their environment.

Danfeng Cai

Danfeng Cai, PhD, combines advanced microscopy, genomics, and proteomics to tease out the functions of protein condensates in cells, with a focus on cancer.

Vivien Thomas PhD Scholars

The  Vivien Thomas Scholars Initiative (VTSI)  is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically Black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. To be considered for the VTSI, you will need to submit a SOPHAS application, VTSI supplementary materials, and all supporting documents (letters, transcripts, and test scores) by December 1, 2024. VTSI applicants are eligible for an application fee waiver , but the fee waiver must be requested by November 15, 2024 and prior to submission of the SOPHAS application.

Per the Collective Bargaining Agreement (CBA) with the JHU PhD Union, the minimum guaranteed 2025-2026 academic year stipend is $50,000 for all PhD students with a 4% increase the following year. Tuition, fees, and medical benefits are provided, including health insurance premiums for PhD student’s children and spouses of international students, depending on visa type. The minimum stipend and tuition coverage is guaranteed for at least the first four years of a BSPH PhD program; specific amounts and the number of years supported, as well as work expectations related to that stipend will vary across departments and funding source. Please refer to the  CBA to review specific benefits, compensation, and other terms.

In the BMB PhD program, all full-time PhD students who remain in good academic standing will receive the above support through the entire duration of the program.

Need-Based Relocation Grants Students who  are admitted to PhD programs at JHU starting in Fall 2023 or beyond can apply to receive a need-based grant to offset the costs of relocating to be able to attend JHU.   These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their PhD program. This is not a merit-based grant. Applications will be evaluated solely based on financial need.  View more information about the need-based relocation grants for PhD students .

Questions about the program? We're happy to help.

Mike Matunis, PhD PhD Program Director

Roza Selimyan , PhD BMB Executive Director for Academic Affairs and Education Programs

Erika Vaitekunas Administrative Specialist

[email protected]

Ph.D. in Biochemistry

General info.

• Faculty working with students: 31
• Students: 55
• Students receiving Financial Aid: 100%
• Part time study available: No
• Application Terms: Fall

Application Deadline: December 2

Meta Kuehn Director of Graduate Studies Department of Biochemistry Box 3711 Duke University Medical Center Durham, NC 27710

Phone: (919) 681-8770

Email: [email protected]

Website:  http://www.biochem.duke.edu

Program Description

Graduate students in the Duke Biochemistry Department engage in faculty-mentored research and graduate coursework that develops an understanding of fundamental concepts in biochemistry, physical biochemistry, as well as the critical analysis of published research. Students choose thesis topics from a wide range of research projects under current study by primary and secondary faculty in the department, which include: enzyme mechanisms, signal transduction, cell cycle control, cytoskeleton structure, prokaryotic and eukaryotic transcription, genetic diseases, glycoproteins, DNA partitioning/segregation, plasma membrane receptors, cellular differentiation, gene regulation, mechanisms of DNA repair and DNA repair defects in tumor biology, processing of DNA, RNA modification, mRNA decay and translational control, multidrug resistance and tolerance, membrane vesicle biogenesis, microbial toxin activity, endocytosis, insertion of proteins into membranes, ion channel structure and function, metalloproteins, post translational modification, drug design, analysis and design of protein and RNA structure, and X-ray crystallography and NMR studies on macromolecular structure and folding. The department is located in a large, research-oriented medical center, which is immediately adjacent to the arts and sciences campus, fostering a productive and collaborative scientific community.

• Biochemistry: PhD Admissions and Enrollment Statistics
• Biochemistry: PhD Completion Rate Statistics
• Biochemistry: PhD Time to Degree Statistics
• Biochemistry: PhD Career Outcomes Statistics

Application Information

Application Terms Available:  Fall

Graduate School Application Requirements See the Application Instructions page for important details about each Graduate School requirement.

• Transcripts: Unofficial transcripts required with application submission; official transcripts required upon admission
• Letters of Recommendation: 3 Required
• Statement of Purpose: Required
• Résumé: Required
• GRE General: Optional
• GRE Subject tests (Biology, Chemistry): Optional This program does not require applicants to provide GRE (Graduate Record Examination) or other graduate entrance exam scores, but does allow you to upload scores if you feel they enhance your application. If you choose to submit test scores, you may enter them on the Test Scores page. If you choose to enter self-reported test scores, official test scores will become a required component of your application.
• English Language Exam: TOEFL, IELTS, or Duolingo English Test required* for applicants whose first language is not English *test waiver may apply for some applicants
• GPA: Undergraduate GPA calculated on 4.0 scale required

Department-Specific Application Requirements (submitted through online application)

Statement of Purpose Guidelines The mission of the Duke University Biochemistry Graduate program is to educate and mentor students from diverse background in the fundamentals of biochemical principles and practice though courses and research by (1) guiding students in their thesis research project, and (2) preparing them for a career in research, education, or other disciplines. The program promotes a commitment to excellence in research scholarship and fosters a spirit of creativity, service, and respect, within an environment that is ethical, inclusive, and diverse.

Writing Sample None required

Additional Components Optional Video Essay: How would a Duke PhD training experience help you achieve your academic and professional goals? Max video length 2 minutes; record externally and provide URL in application.

We strongly encourage you to review additional department-specific application guidance from the program to which you are applying: Departmental Application Guidance

List of Graduate School Programs and Degrees

Biochemistry, Cellular and Molecular Biology, PhD

School of medicine, fields of study and research.

The Departments of Biological Chemistry, Biophysics and Biophysical Chemistry, Cell Biology, Molecular Biology and Genetics, Neuroscience, Pharmacology and Molecular Sciences, and Physiology jointly provide a program of study leading to the PhD degree. The objective of the Biochemistry, Cellular and Molecular Biology (BCMB) program is to provide students with the breadth of knowledge and research training that will prepare them for their own independent and productive careers.  Strong emphasis is placed on molecular and cellular approaches to fundamental problems in research areas covered by the participating departments. The focus on a broad background in basic science for all students promotes the study of problems of biomedical importance from a mechanistic perspective. The multi-departmental nature of the program fosters a collaborative environment that is ideal for graduate training, and promotes the extensive interaction and collegiality that is a hallmark of Johns Hopkins science.

Training includes a rigorous and broad-based first year curriculum, research rotations in three different laboratories, a proposal-based qualifying exam (along with a proposal-writing workshop), and thesis research in the laboratory of one of more than 100 participating faculty members. Once they join a laboratory, students establish a departmental affiliation and participate in their departmental journal clubs and seminars. At the same time, students continue to maintain their BCMB identity and affiliation in a number of ways. These include attending the annual BCMB retreat, assisting in the recruitment of new students, hosting a one-day symposium with outside speakers, and participating in a monthly student research colloquium (organized by BCMB students for their peers). There is no formal teaching requirement for BCMB students, but many opportunities for gaining teaching experience are available.

Due to the multidisciplinary focus of the BCMB program, students are very well prepared for the full breadth of modern biomedical science careers. Greater than 90% of students in the BCMB program complete the PhD degree. The average time to degree over the last 10 years is 5.7 years. On average, BCMB students publish 4 research papers with 2 as first author.

To date, there are over 750 BCMB graduates; many hold leadership positions in academia, industry, government and other sectors. At least 97% of graduates in the last 5 years are following career paths in laboratory research, medicine, or research-related careers.

The lecture halls and the research laboratories are located in the Wood Basic Science Building and the adjoining Physiology, Biophysics, Hunterian, Preclinical Teaching, Ross Building, Miller Research, and Rangos Buildings. Modern equipment and state-of-the-art core facilities are available to students. This includes electron microscopes, confocal microscopes, nuclear magnetic resonance spectrometers, mass spectrometers, HPLC and gas-liquid chromatographs, protein sequencers, peptide synthesizers, oligonucleotide synthesizers, X-ray diffraction equipment, as well as instrumentation for microarray analysis and deep sequencing.

Collaborative Facilities

All seven departments enjoy excellent working relationships with other departments in the medical school and with departments at the Homewood campus.

Financial Support

All students accepted into the BCMB program receive full tuition, health and dental insurance, and stipend support for the duration of their studies.

Admission Requirements

Candidates for admission should show a strong academic foundation with coursework in biology, chemistry, physical sciences, and quantitative analysis. A bachelor's degree from a qualified college or university will be required for matriculation.

The BCMB website has up-to-date information on “ How to Apply ” and “ Frequently Asked Questions .” For questions not addressed on these pages, please email [email protected] .

Program Requirements

Students must successfully complete all eight courses of the “Foundations of Modern Biology” core curriculum, and the Core Discussion course in the first three quarters of year one. During the fourth quarter, students complete two electives, the “BCMB Responsible Conduct in Research” course, and the “Method, Logic and Experimental Design” course.  Three additional elective courses are required in the advanced years of study in addition to the BCMB "Seminar" and "Writing a Paper" courses.

An oral examination, conducted as prescribed by the Doctor of Philosophy Board of the University, must be completed before the end of the second year. Subsequently, students are required to participate in annual thesis committee meetings as they work on their thesis project, to review research progress and discuss plans for the next year.​​​

Usually in year five, the student's thesis committee agrees that the student is nearing completion of their research and will be ready to write a dissertation. The student's advisor and one other member from their thesis committee will read and approve the dissertation. The student must present a public seminar on their completed thesis research.

The BCMB program recognizes the importance of students’ professional development and career exploration during their graduate studies. In order to better integrate professional development into the training, the BCMB program is formally participating in the “OPTIONS” Career Curriculum offered by the Professional Development and Career Office (PDCO). During the first two years, students attend workshops and panel discussions facilitated by professionals in different fields discussing their career paths, what each profession entails, how the graduate training relates to the required skills and knowledge, and what additional skills would be useful. Several of the speakers are BCMB program alumni, and offer to follow-up with interested students, fostering a sense of community. In year 3, students join an OPTIONS Career Community which comprises six monthly workshops that provide exposure to careers and skills assessment (e.g., Academic Research; Business and Finance; Biotech and Pharma; Science Communication and Policy; Academic Teaching; etc.). In year 4 and beyond, students meet with a career coach to create a plan for gaining more career-relevant experiences and/or participate in an internship.

The first year required courses are as follows:

Courses are detailed under the course descriptions listed in the entries of the Departments of Biological Chemistry, Biophysics and Biophysical Chemistry, Cell Biology, Molecular Biology and Genetics, Neuroscience, Pharmacology and Molecular Sciences, and Physiology, or as Interdepartmental.

Graduate Programs

• Biochem & MolBiophysics PhD

Biochemistry and Molecular Biophysics PhD

The Biochemistry and Molecular Biophysics PhD Program ranks in the top 10 nationally and represents a traditional strength in the Chemistry and Biochemistry Department at UCSD. The goal of the program is to prepare students for careers in the biochemical sciences as researchers and educators by expanding their knowledge of structural biology, protein, RNA, and lipid biochemistry, experimental and computational biophysics, and systems biology while developing their ability for critical analysis, creativity, and independent study. A high graduation rate in an average of just over five years can be attributed to the quality of applicants admitted, the flexibility of our program of study, the opportunity for students to begin research in the first year, and the affordability of education made possible by our generous financial support policies.

Program Overview

Programs of study are tailored to the needs of individual students, based on their prior training and research interests. However, progress to degree is generally similar for all students. During the first year, students take courses, begin their teaching apprenticeships, choose research advisors, and embark on their thesis research; students whose native language is not English must pass an English proficiency examination. Beginning the first summer, the emphasis is on research, although courses of special interest may be taken throughout a student's residency. At the end of their first year, students choose the departmental members of their thesis committee and begin to prepare a written research proposal. During their second year, they complete their research proposal and defend it orally. In the third year, students advance to candidacy for the doctorate by defending the topic, preliminary findings, and future research plans for their dissertation. Subsequent years focus on thesis research and writing the dissertation. Most students graduate during their fifth year.

Research Opportunities

Research opportunities for graduate students are comprehensive and interdisciplinary, spanning biochemistry; biophysics; structural biology, protein, RNA, and lipid biochemistry, experimental and computational biophysics, and systems biology. Please refer to the faculty pages for full descriptions of the on-going research of faculty in the Biochemistry and Molecular Biophysics PhD Program. State-of-the-art facilities and laboratories support these research programs.

UCSD is a thriving community that stretches across campus with opportunities for research and collaborations among a large number of faculty in the Division of Biology, the Skaggs School of Pharmacy and Pharmaceutical Sciences, the School of Medicine, the La Jolla Institute of Immunology, the Salk Institute, and many others.

Special Training Programs

Interdisciplinary research and collaboration at UCSD is enhanced through a variety of training grants. These programs provide financial support for exceptional graduate and postdoctoral scholars and also unite researchers from across campus and throughout the La Jolla research community in special seminars, retreats, and courses. Doctoral students usually apply for training grants in their second year.

• Molecular Biophysics Training Grant
• Contemporary Approaches to Cancer Cell Signaling and Communication
• Interfaces Graduate Training Program
• Molecular Pharmacology Training Program

Teaching apprenticeships are a vital and integral part of graduate student training, and four quarters of teaching are normally required. See the Teaching Assistants page to apply. Students can gain experience teaching both discussion and laboratory sections. Excellence in teaching is stressed, and the department provides a thorough training program covering both fundamentals and special techniques for effective instruction. Further training is provided by the Teaching and Learning Commons on campus. Performance is evaluated every quarter, and awards are bestowed quarterly for outstanding teaching performance.

Financial Support

Students in good academic standing receive a 12-month stipend; fees and tuition are also provided. Support packages come from a variety of sources, including teaching and research assistantships, training grants, fellowships, and awards. Special fellowships are awarded to outstanding students based on their admission files. See Ph.D. Program Support Policy for more information.

Health and Dental Plan

A primary health care program, major medical plan, and dental plan are among the benefits provided by the University's registration fee (see Graduate Student Health Insurance Program, GSHIP) . Minor illnesses and injuries can usually be treated at the Student Health Center . Counseling is provided free of charge through Counseling and Psychological Services .

Creative, bright, and motivated students from diverse backgrounds are encouraged to apply. We admit for Fall quarter entrance only. Generally, the application will open in September and close is in December. The Admissions Committee reviews files individually and in comparison to others, and invitations to interview are made around January. For those invited, in-person interviews will be on campus in either February or March. Those who receive the official admissions offer from t he Division of Graduate Education and Postdoctoral Affairs ( GEPA ) have until April 15th to make a decision.

PostGraduate Placement

Graduates typically obtain jobs in academia or in the biotech/pharmaceutical industry. La Jolla is home to the third largest Biotech/Pharmaceutical industry mecca. Many of our alumni stay in San Diego and obtain positions in one of the over 300 companies that are located near UCSD. During their PhD, students can take advantage of the many internships that are available at these companies. A large proportion of our graduates attain postdoctoral research positions in leading academic institutions. The Biochemistry and Molecular Biophysics Program provides career advising throughout the PhD. UCSD's Career Services Center and the Physical Sciences Student Success Center provides many resources for students, including the chance to videotape yourself in a mock interview!

• BMB Degree Requirements
• BMB Financial Support
• Chemistry PhD
• Masters Program
• Joint Doctoral Program
• Teaching Assistants
• Student Spotlight
• Graduate Events
• Course Offerings
• Room Requests

Course Descriptions

A Degree Built By You

You’ll customize your plan of study with help from faculty advisors and your research mentors, choosing from a broad list of available courses in the  biomedical science graduate curriculum . In the first 2 years of study, you are required to complete a minimum of 24 hours of graded coursework (including a total of 8 units of Independent Study Research), as well as at least 6 hours of ungraded coursework (typically met with the  RCR training .)  Note  that students coming into the program from the Medical Scientist Training Program (MSTP) or with an MA/MS degree, may be exempt from the minimum graded coursework requirement.

Your Faculty Advisory Committee and Research Mentors play an important role in your graduate student career development.

Faculty Advisory Committee:

• Customizes your plan of study to include basic and advanced courses in and around your research focus.
• Aids in formulating research objectives and thesis topics for success in the Preliminary Qualifying Exam.
• Guides annual progress on your dissertation.
• Advises on the skills and experience you’ll need to reach your professional goals.

Research Mentors:

• Customize your plan of study with day-to-day troubleshooting and by developing a long-term thesis project.
• Inform you of different career opportunities.
• Promote your presentations at local, national, and international meetings and workshops.
• Help set and revise short- and long-term goals.

" I chose Duke Biochemistry for its premier structure-function research, welcoming faculty and amiable atmosphere "  —Kalina Tsolova, 2021 Matriculant

Fall and spring semester courses, of particular interest to Biochemistry PhD students, are described below. Courses offered by other departments in the Basic Sciences can be found on the School of Medicine Biomedical Graduate Education  website  and in the Biochemistry  Graduate Student Handbook .

Instructor:  DGS-Kuehn Fall and Spring Semesters (Years 1 and 2) In year one, you will rotate through and conduct faculty/mentor-approved research in different Biochemistry labs. If you’re in your 2 nd  year, you’ll be in a thesis lab. Students are expected to be performing research when they are not in class, attend departmental seminars, keep up-to-date lab notebooks, and present oral and written summaries of their ongoing research, as appropriate for each laboratory.

Instructor:  Richardsons Not Offered Spring Semester 2023 You will learn how to get the most out of experimental and computational 3D structures including:

• 3D Molecular Literacy: computer and physical molecular models of proteins and nucleic acids, worksheets, and hands-on exploration.
• Data bases and the data itself: gaining familiarity with the Protein Data Bank (PDB); the Electron Density Server (EDS); and the peculiarities, caveats, and reliabilities of different categories of molecular data.
• Model building in structural biology
• Molprobity and all-atom contact analysis and methodologies for multiple conformations, ensembles, and mobility

Projects include: interactive 3D illustrations about macromolecules using kinemages or other molecular graphics programs, often with short, non-interactive introduction. 

Instructors : Lee, Bennett, Kuehn, Rajagopal, and Yang 2 nd  ½ Spring  Semester This course will highlight modern topics regarding biological membranes and membrane proteins that are important for human physiology and disease. Topics include:

• Structure and dynamics of biological membranes
• Structure, and function of membrane proteins that play critical roles in cell signaling
• Diseases related to dysfunction of membrane and membrane proteins
• Current efforts on drug discovery

Major techniques used in membrane research will also be covered. The format will be a combination of lectures and discussions of primary literature. Students will be evaluated based on their class participation and performance at the final presentations. 

Instructor:  Beese 1 st  ½ Fall Semester Structure of Macromolecules – This class teaches the principles of modern structural biology including protein-nucleic acid recognition, enzymatic reactions, viruses, immunoglobulins, signal transduction, and structure-based drug design—described in terms of the atomic properties of biological macromolecules. There will be discussions on the methods of structure determination, with particular emphasis on macromolecular X-ray crystallography, Nuclear Magnetic Resonance (NMR) methods, homology modeling, and bioinformatics. You will use molecular graphics tutorials and Internet databases to view and analyze structures.

Instructor:  Zhou 2 nd ½ Fall Semester Molecular Biology I – As a continuation of BCH 658, this introductory course teaches how to use quantitative methods to understand biological structure and function. You’ll study the:

• Structure/function analysis of proteins as enzymes
• Kinetics of binding
• Catalysis and allostery
• Protein folding
• Design protein-protein interactions

Instructor:  Schumacher Not Offered Spring Semester 2023 A major emphasis of this course is on the reading and in-depth discussion of primary research papers. You will explore how new nucleic acid biology concepts have been developed and what types of experiments have made advances possible. Each course section will consist of background lecture material presented by the faculty member, followed by several sessions to discuss paradigm papers selected by the faculty. You will do oral presentations using figures from the papers and explanatory background as needed and prepare an 8-10 page research paper or an oral talk to demonstrate proficiency in the topics of DNA and Genome Stability—Chromatin and chromosome structure, replication, repair, genetic recombination, mutation, and chromosome rearrangement.

Instructor:  Meyer Spring Semester This course explores various aspects of RNA biology and function. Topics include:

• Translation
• RNA: Protein interactions
• Non-coding RNAs
• RNA modifications
• Viral RNA regulation
• RNA structure-function relationships
• RNA-targeted drug discovery

You will also learn about major techniques in RNA research, including in vitro and in vivo methods for understanding global RNA regulation. The format will be a combination of weekly lectures and discussions of primary scientific literature. You will be evaluated on your participation and performance during in-class presentations and will write a short mock research grant on a topic of your choosing.

Instructor:  Oas Fall Semester You’ll survey the theory and application of biological techniques—based on physical principles—to understand their application in primary scientific literature. Methods covered include:

• Optical Spectroscopy
• Fluorescence
• Light Microscopy
• Electron Microscopy
• Hydrodynamics
• Kinetic Methods
• Electrophysiology
• Magnetic Resonance
• Mass Spectrometry

Lectures on each method are presented by an expert. 

Instructor:  Bartesaghi Fall Semester This Advanced Topics course will focus on the image analysis aspects of cryo-electron microscopy (EM), including image enhancement, reconstruction, classification, and movie processing used to determine the high-resolution structure of proteins from cryo-EM images. The course will provide an overview of the cryo-EM structure determination pipeline, focusing primarily on the data analysis aspects of the technique and covering the full breadth of reconstruction strategies.

Prerequisites: Students should have a background in either protein structure/molecular biology or computer vision/image processing.

Instructor:  Al-Hashimi Spring Semester  (alternating years with BCH 696) Aimed for students with some familiarity in high-resolution NMR, you will deepen your understanding of how NMR experiments actually work and be introduced to quantum mechanical tools that help explain pulse sequences. You’ll study advanced biomolecular NMR experiments that enable the structural and dynamic characterization of biomolecules. Roughly half the course follows online lectures that accompany the textbook, with class meetings emphasizing concepts, group discussion, and problem-solving.  Instructor consent is required.

Instructor:  Schumacher and Staff Spring Semester (alternating years with BCH 695) This course teaches the theoretical and practical principles of macromolecular X-ray crystallography. Topics include:

• Crystal symmetry
• Space group theory and determination
• Diffraction theory
• Practical understanding of crystallization
• X-ray intensity data collection and data processing
• Phase determination
• Refinement and model validation

Instructor consent is required.

Instructor:  Brennan Fall/Spring Semester This course is a requirement of all first-, second-, and third-year biochemistry graduate students to learn how to clearly and concisely present the background, data, conclusions, and future prospects of your research in both oral and written formats. First-year students will present their rotation projects. Second and third-year students will annually present their research. All students will provide feedback so each presenter can improve presentation content and style.

Instructor:  Kuehn Fall Semester This is a discussion-based course that includes introductory lectures and the critical reading of select topics in Biochemistry literature. Topics and instructors are announced each semester. 

Instructor:  Noor 1 st  ½ Fall Semester This course is a great introduction to understanding the graduate student mindset. Weekly talks and Q&As will focus on choosing a thesis advisor, writing a grant proposal, the scientific manuscript peer review processes, and other topics related to transitioning to and being successful in graduate school. 

Instructor:  Oas Fall and Spring Semester Each week, a student will present a paper on their research. Attendance is open to all graduate and postdoctoral students and faculty who have an interest in structural biology. All SBB certificate students are required to attend.

Instructor:  McCafferty Spring Semester This course covers:

• Basic enzymology
• Mechanisms of enzymatic reactions
• Oxidoreductases
• C1 chemistry
• Carbon-carbon bond formation
• Carboxylation/decarboxylation
• Pyridoxal enzymes
• Thiamine enzymes

Instructor:  Caron First 1/3 Spring Semester This module covers the basic mechanism of signal transduction through G protein-coupled receptors (GPCR) and how they control a wide array of biological functions—from vision to reproduction—and how they are the largest targets of therapeutic interventions. You will study how new concepts of signal transduction mechanisms are leading to the development of new and improved therapies for various disorders.

Instructor:  McDonnell Mid 1/3 Spring Semester This module covers how ion channels and intracellular nuclear receptors control cellular functions, mediated through transcription or calcium signaling, to regulate physiological processes in health and disease.

Instructor:  Pendergast Last 1/3 Spring Semester This module focuses on signaling pathways induced by extracellular factors that regulate growth, survival, and development—and their deregulation in diseases including cancer. Topics include those regulated by ligand-activated Receptor Tyrosine Kinases, Wnt/beta-catenin signaling, Notch signaling, and Hedgehog signaling.

PhD Program

Be part of innovative research and mentored by leading experts in the field.

Every student must publish at least one first author research paper prior to graduation

Our department offers a PhD in Biochemistry and Molecular & Cellular Biology. The program is designed for students interested in molecular mechanisms, proteomics, and biophysics. The department accepts up to five PhD students each year.

What You’ll Learn

Your training as a doctoral candidate in the Department of Biochemistry and Molecular & Cellular Biology at GUMC will be multifaceted. Our program has developed a solid educational platform designed not only to challenge the candidate, but also to provide supportive mentorship in order to aid the candidate as they develop their own skillsets and become a scientific leader.

Thesis mentors are available from a variety of backgrounds spanning stem cell biology and human tissue engineering to traditional biochemical analytics, with an optional focus on informatics- or systems-based medicine. Each faculty mentor represents a diverse component of the biomedical scientific enterprise and each has dedicated their career to enhancing biomedical science through independent research and doctoral candidate training.

Graduates of the program have gone on to successful careers in academia, industry, medicine, law, and finance. Contact the program if you would like more information.

Learn about the application process.

Browse our required courses.

Course Schedule

Find an example course schedule.

Student Resources

Many resources are available for current students.

Career Services

Prepare for the next step in your career.

Questions about the program or application process? Email us!

• Degree Programs

PhD in Biochemistry

Application information.

Students enter the Biochemistry PhD program by applying to the Biomedical Sciences Training Program (BSTP) .  Ideal candidates will possess strong grades and have completed undergraduate courses in biology, mathematics, physics, and chemistry.

To apply students should submit

• a completed application form,
• three letters of recommendation,
• official undergraduate transcript(s), and
• when applicable, official TOEFL scores.

If you wish to apply only to the Biochemistry PhD program, select Biochemistry in the "Preferred Program of Interest (PPI)" menu. Please read about PPIs in  the BSTP's FAQ page .

Online applications are submitted through the site for the School of Graduate Studies. Please select Biomedical Sciences Training Program in the "Academic Program" menu once you  start the application .

Biochemistry, PHD

On this page:.

At a Glance: program details

• Location: Tempe campus
• Second Language Requirement: No

Program Description

Degree Awarded: PHD Biochemistry

The PhD program in biochemistry through the School of Molecular Sciences provides students with the training they need to solve biological problems at the molecular scale and to be successful, independent scientists who can challenge current societal issues.

Students earning a doctoral degree in biochemistry are fully trained in all fundamental aspects of the discipline. Most also choose to join transdisciplinary teams that work on larger, mission-based contemporary problems in areas such as:

• biogeochemistry
• energy and sustainability
• fundamental chemical biology
• materials and nanoscience
• medicine and health
• structure function and dynamics

Students should visit the prospective student site to learn more about this doctoral program and the graduate research page to learn more about the advanced transdisciplinary research being conducted in the School of Molecular Sciences.

Degree Requirements

Curriculum plan options.

• 84 credit hours, a written comprehensive exam, an oral comprehensive exam, a prospectus and a dissertation

six graduate-level courses (12--18) enrollment in a BCH 501/CHM 501 seminar (8) BCH 792 Research (46--52) BCH 799 Dissertation (12)

Additional Curriculum Information The program consists of coursework and seminars selected by the student in consultation with the student's supervisory committee and based on the student's area of research. Qualifying exams consisting of a written comprehensive exam, an oral comprehensive exam and a prospectus are required to advance to candidacy. Students must successfully defend their dissertation during a public final oral defense. Students must also maintain a minimum GPA of 3.00 (scale is 4.00 = "A") or better.

Admission Requirements

Applicants must fulfill the requirements of both the Graduate College and The College of Liberal Arts and Sciences.

Applicants are eligible to apply to the program if they have earned a bachelor's or master's degree in chemistry, biochemistry or a closely related field from a regionally accredited institution.

Applicants must have a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in the last 60 hours of their first bachelor's degree program.

All applicants must submit:

• graduate admission application and application fee
• official transcripts
• personal statement
• three letters of recommendation
• proof of English proficiency

Additional Application Information An applicant whose native language is not English must provide proof of English proficiency regardless of their current residency.

In addition to the admission application, applicants must complete an online supplemental application to the School of Molecular Sciences. The supplemental application is available 24 to 48 hours after submission of the admission application. The personal statement and contact information for the letters of recommendation are submitted as part of the supplemental application. Information about the supplemental application can be found on the department's website. Applications lacking a supplemental application are not reviewed.

Next Steps to attend ASU

Learn about our programs, apply to a program, visit our campus, application deadlines, career opportunities.

The doctoral program in biochemistry prepares students for professional careers in industry, government and academia. Professionals with training achieved in pursuit of a graduate degree in chemistry or biochemistry have opportunities in five general areas:

• academia (high school and higher education)
• entrepreneurship (consulting, startups)
• government (research, policy)
• industry (research and development, quality control)
• nonprofit (policy, public education)

In addition to specialized technical skills, graduates possess high-demand skills like critical thinking, teamwork and collaboration, time management and many others.

Some career examples include:

• chemistry lecturer
• drug discovery scientist
• government scientist
• pharmacology scientist
• research and development scientist
• research group leader
• science consultant
• university professor

The American Chemical Society also provides helpful resources and a more exhaustive list of possible careers on their website at https://www.acs.org/careers/chemical-sciences.html .

Program Contact Information

If you have questions related to admission, please click here to request information and an admission specialist will reach out to you directly. For questions regarding faculty or courses, please use the contact information below.

• [email protected]
• 480/965-4664

The Biochemistry and Molecular Biology doctorate degree program at IU School of Medicine typically takes five years to complete. The program includes two years of course work and three years of conducting original research full-time, culminating in a PhD thesis. This program prepares graduates for careers in academic research and teaching or in the business and industry side of bioscience.

Program Requirements

Graduation requirements.

• A minimum of 30 credit hours of didactic coursework
• A minimum of 45 credit hours of research (B855)
• A minimum of 90 credit hours consisting of required and elective courses
• A scholarly thesis based on approved original research with oversight from a faculty mentor. Potential research topics include structural biology, signal transduction, drug development, regulation of gene expression, cancer biology and metabolic disorders.
• Minimum 3.0 GPA and a B- or better in all PhD coursework

Curriculum Year One

Biochemistry PhD students must take at least two (of the available eight) two-credit Biochemistry core courses (BIOC-B811, GRDM-G805, G807, G817, G848, G852, G825, G826), which are offered among spring electives for the Indiana Biomedical Gateway Program (IBGP) or in the fall. These may be taken in year one or later years.

• GRDM-G715 Biomed I – Biomedical Science I – Biochemical Basis of Biological Processes*: 2 credits
• GRDM-G716 Biomed II – Biomedical Science II – Molecular Biology and Genetics*: 2 credits
• GRDM-G717 Biomed III – Biomedical Science III – Cellular Basis of Systems Biology*: 2 credits
• GRDM-G505 Responsible Conduct of Research*: 1 credit
• GRDM-G700 Translating Foundational Science to Contemporary Knowledge*: 1 credit
• GRDM-G702 Entering Biomedical Science Research*: 1 credit
• GRDM-G718 Research in Biomedical Science (first rotation): 2 credits

Biochemistry Core Electives

• GRDM-G805 Diabetes and Obesity: 2 credits
• GRDM-G825 Advanced Topics in Molecular Biology: 2 credits
• GRDM-G826 Synthetic Biology and Biologics: 2 credits

Research Communication (one of the following)

• COMM-C534 Distilling Your Message*: 1 credit
• COMM-C533 Improvisation for Scientists*: 1 credit
• ENG-W533 Science Writing for Public Readers*: 1 credit
• GRDM-G507 Reagent Validation as a Means for Enhanced Research Reproducibility*: 1 credit
• GRDM-G718 Research in Biomedical Science Rotations 2 and 3: 4 credits
• Indiana Biomedical Gateway Program open enrollment electives
• GRDM-G817 Molecular basis of cell structure and function: 2 credits
• GRDM-G852 Concepts of cancer biology: 2 credits
• GRDM-G807 Structural and chemical biology: 2 credits
• GRDM-G848 Bioinformatics applications to proteomics and genomics: 2 credits

Biochemistry Course Descriptions

Curriculum Year Two

Fall- Total of 10 credit hours

• GRDM-G855 Experimental Design and Research Biostatistics*: 1 credit
• BIOC- B890 Seminar: 1 credit
• Major course requirements: variable
• Minor course requirements: variable
• BIOC-B855 Research: variable

Spring- Total of 10 credit hours

• BIOC-B803 Advanced Biochemistry: 1 credit

Curriculum Years Three through Five

During years 3 through 5, the student will take didactic courses if needed to fulfill either requirements for the biochemistry major or their chosen minor. In year 3, the student will typically register for a total of 10 cr. hours each fall and spring, including 1 cr. hour of BIOC-B890. 4 credit hours of B890 are required before advancing to candidacy.   The student will advance to candidacy upon completion of both major and minor coursework and having reached a total of 30 or more didactic hours of coursework as well as successful defense of their thesis proposal which serves as the students qualifying examination.

Work will continue in the field of the candidate’s thesis. Emphasis will be on the ability to pursue research with relative independence and responsibility.

Curriculum for Dual Degree (MD/PhD) Program

MD/PhD students are not required to take GRDM-G715-717 but are expected to perform lab rotations during summer breaks from medical school classes. MD/PhD students must take two or more of the two-credit Biochemistry core courses (GRDM-G805, G807, G817, G852, G825, G826, G848) along with other courses required of IBMG and Biochemistry and Molecular Biology PhD students (GRDM-G505, G655, G855, BIOC-B803 and B890 [4x]).

Thesis Requirement

After a laboratory for thesis research is chosen, an advisory committee consisting of at least three faculty from Biochemistry and Molecular Biology and one external faculty member is formed with the approval of the thesis advisor and departmental chairperson. Upon advancement to candidacy, a thesis research committee is similarly formed but may consist of different faculty.

Financial Aid

 PhD students with eligibility to work in the U.S. receive a competitive annual stipend without a Teaching Assistant requirement ($29,000 for the 2020-2021 academic year) as well as tuition scholarships and health and dental insurance. Find additional financial aid information from the IUPUI Office of Student Financial Services .

• Interesting for you
• My settings

Biochemistry in United States

Why Study Biochemistry in United States

• Studying Biochemistry in United States is a great choice, as there are 77 universities that offer PhD degrees on our portal.
• Over 957,000 international students choose United States for their studies, which suggests you’ll enjoy a vibrant and culturally diverse learning experience and make friends from all over the world.
• We counted 127 affordable PhD degrees in United States , allowing you to access quality higher education without breaking the bank. Moreover, there are 299 available scholarships you can apply to.

98  Biochemistry PhDs in United States

Chemical and Biochemical Engineering The Chemical and Biochemical Engineering PhD program from the University of Maryland Baltimore County (UMBC)... University of Maryland Baltimore County (UMBC) Baltimore Highlands, Maryland, United States

Biochemistry This Biochemistry PhD program from the University of Maryland Baltimore County (UMBC) offers a course of... University of Maryland Baltimore County (UMBC) Baltimore Highlands, Maryland, United States

Biochemistry The Biochemistry PhD program at Medical College of Wisconsin will expose you to state-of-the-art facilities... Medical College of Wisconsin Milwaukee, Wisconsin, United States

Study in United States

The United States is home to some of the most prestigious universities and colleges in the world. With over 150 universities featured in international rankings, the U.S. has some of the best business schools, medical schools, and engineering schools. Universities and colleges in the U.S. are well known for academic flexibility and ways to customize your study experience with optional studies and extracurricular activities. Depending on where you will be studying, you will be able to visit iconic places like the Statue of Liberty, Empire State Building, Goldengate Bridge, The Grand Canyon, Mount Rushmore, Disney's Magic Kingdom Park, and much more.

Is United States the right place for you?

Take the test and find out which country is your best fit.

Explore your Biochemistry degree

Biochemistry degrees teach students about the chemical processes that take place inside living organisms. Biochemistry is usually split into three branches of Science: Molecular Genetics, Protein Science, and Metabolism. After graduation, Biochemistry students can find jobs as toxicologists, forensic or analytical scientists, clinical research associate, clinical biochemists, and more.

Is Biochemistry the best for you?

Take the test and find out if Biochemistry is the right path for you.

Biochemistry Scholarships and Financial Aid in United States

Available Options

Go to your profile page to get personalised recommendations!

College of Liberal Arts & Sciences

School of Molecular & Cellular Biology

• Meet the MCB Community
• Instruction & Advising
• MCB Academic Departments
• Supporting Offices & Resources
• Health & Wellbeing Resources
• Job Opportunities in MCB
• Undergraduate Admissions
• Master's Program Admissions
• PhD Program Admissions
• Major in MCB
• Major in Neuroscience
• Major in Biochemistry
• Pre-med & pre-health guide
• Master's Program
• Prospective students
• Current students
• School of MCB Convocation
• Biochemistry
• Cell & Developmental Biology
• Microbiology
• Molecular & Integrative Physiology
• Explore Research in MCB
• Research Collaborations & Partners
• Undergraduate Research
• Research Facilities & Resources
• In the News
• MCB Convocation
• Seminars and Events
• Graduate Research Retreat
• Awards and Honors
• In Memoriam
• Media contacts in MCB
• Diversity, Equity & Inclusion in MCB
• MCB DEI Committee
• Campus Diversity Offices and Resources
• Alumni & Friends Community
• Mentorship Program
• Share Your News & Update Contact Info
• Giving opportunities
• Make an Online Gift
• School Leadership & Support Staff
• Advising & Instruction Team
• All MCB Faculty

Department of Biochemistry

Pursue a PhD in Biochemistry

Request information    Degree requirements   Apply Core courses   Advanced Electives

We train graduate students to become skilled and creative biochemists and molecular biologists. We have outstanding resources in our internationally recognized faculty, graduate students, and research facilities. Our graduate program not only allows our research students to learn the latest techniques, but also fosters their development as independent scientists. With a PhD in Biochemistry from UIUC, you will be well-prepared to launch a career in academia or industry. Our alumni are leaders in industrial laboratories, research institutes, and government agencies, as well as in teaching, research, and administration in colleges and universities.

As a PhD student in the Department of Biochemistry, you will pursue research alongside brilliant faculty and graduate students while advancing the frontiers of this exciting field. Our labs encompass a variety of areas, including biological chemistry and molecular biology: physical approaches to the structure and function of proteins, nucleic acids, and carbohydrates, including structural biology (Cryo-Em, x-ray crystallography, etc.); genomics, enzymology; membrane biochemistry and protein-lipid interactions; protein-nucleic acid interactions; molecular biological approaches to gene organization and expression; immunology; microbial physiology, and signal transduction. We collaborate with peers at units across campus and around the world.

At the U of I we boast world-class facilities, equipment, and technology . The department is a close-knit community which holds numerous events throughout the year, such as an annual research conference and graduate research seminar series.

Title Meet Yiquan

When he's not studying SARS-CoV-2 or influenza, Yiquan Wang, a biochemistry PhD student and member of Nicholas Wu's lab, can be found creating works of art. Yiquan shares how he's bridging his love for art and biochemistry to help people better understand virology. 

Title State-of-the-art facilities

The Department of Biochemistry contains all of the equipment appropriate for modern biochemical research. Specialized facilities available within the department include equipment for large scale growth and processing of animal cells as well as state-of-the-art S-ray diffraction equipment. Other supporting facilities include specialized labs for recording infrared, ultraviolet, fluorescence, electron spin resonance, mass, nuclear magnetic resonance, and optical rotatory dispersion spectra. Electronic, machine, photographic, and glass-blowing shops are also available.

The campus also provides many state-of-the-art core facilities, from the Beckman Institute to the Woese Institute for Genomic Biology .

Research collaborations   Research facilities & resources 

The Department of Biochemistry is part of the School of Molecular & Cellular Biology. Prospective Biochemistry PhD students should apply directly to the MCB PhD Program .

During your first semester, you will rotate through three laboratories to learn experimental techniques and the information you need to choose a research project. You will then select a research adviser and formally join the graduate program by the spring semester of your first year.

Financial support

All students accepted into the PhD program receive year-round financial support through fellowships, traineeships, or graduate assistantships, which provide a generous stipend as well as a full tuition and partial fee waiver. The department guarantees financial support to all students as long as they make satisfactory progress toward their PhD. Students are also encouraged to apply for extramural pre-doctoral fellowships for which they may be eligible, such as the National Science Foundation, Howard Hughes Medical Institute, Ford Foundation, and other fellowships.

MCB PhD Programs

Shawna M. Smith, MCB Graduate Program Coordinator Lori Raetzman, Associate Director of MCB Graduate Program [email protected] ; 217-333-1737

Graduate student affairs: [email protected]

Satish Nair, Department Head and Gregorio Weber Chair [email protected] ; 217-333-3945

Kai Zhang, Associate Professor of Biochemistry and Director of Graduate Studies; Associate Head, Department of Biochemistry [email protected]

• Alumni and Giving
• International Programs
• Undergraduate
• Undergraduate Curriculum
• Undergraduate Advising Resources
• Frequently Asked Questions
• Spend A Day At State
• Graduate Student Resources

Ph.D. in Biochemistry

• IBMP (Interdisciplinary Biochemistry Masters Progam)
• Accelerated Bachelor’s/Master’s Degree
• Graduate Admissions and Guidelines
• Professional Development
• Graduate Students
• Associated Faculty
• Adjunct Faculty
• Give Now 

Students entering into our Ph.D. program generally have a B.S. in biochemistry, chemistry, biology, genetics, or related physical or biological science. During the first year in residence, students are advised by a graduate advisory committee and participate in coursework, teaching, and laboratory rotations.

Core Courses

The core courses in the biochemistry Ph.D. program include:

• BCH 590: Special Topics: Experimental Design and Application
• BCH 701: Macromolecular Structure and Metabolism
• BCH 703: Transcription, Translation, and Genome Dynamics
• BCH 705: Signal Transduction and Cellular Regulation
• BCH 801 and 810: Departmental Seminar (Thursdays) and Graduate Seminar (Fridays)

Following the core coursework, students may enroll in up to two advanced courses, chosen by the student based on recommendations from their thesis advisor, designed to support their thesis project.  Advanced topics range from biophysical chemistry, advanced molecular biology, metabolism, bioinformatics, or several topics offered through other departments on campus. Students usually complete coursework is completed by the end of the second year. In addition, students gain teaching experience by participating as teaching assistants for one or more semesters. Students also attend weekly departmental seminars presented by eminent visiting scientists.

Graduate Support

We support doctoral students through teaching assistantships, research assistantships, or faculty grants. In addition to stipend support, the department covers tuition for doctoral students and medical benefits through the Graduate Student Support Plan.

Application and Admission Timeline

Review of applications begins in November, and applicants are strongly encouraged to apply well before the December 1, 2023 target date for fall admission, although applications will be accepted until January 15, 2024 .  Well-qualified applicants will be contacted in late December-early January. Due to ongoing health and travel concerns, the initial phases of the process will be virtual. Campus visits will be mid-Spring semester.

Incoming Students

Students are advised to arrive in Raleigh the first week of August to allow time to acclimate (exploring your new neighborhood, getting your North Carolina driver’s license, etc). The week before classes start (typically the second week of August), incoming students will meet with the director of graduate programs and the department head for initial advising of coursework. During this week, you will also complete paperwork, obtain your student ID, and secure a parking permit (if needed).

Doctoral students participate in two laboratory rotations, each approximately eight weeks in duration, which allow the student to choose a thesis advisor through knowledge of the laboratory environment.  You will have time to meet with current graduate students and faculty to learn more about potential rotation projects during orientation week. Students will select their first laboratory rotation at the end of orientation week and their second rotation four weeks into the semester. The first rotation continues until fall break, and the second rotation ends before finals week.

Selecting a Laboratory

At the end of the first semester, the student will select a thesis advisor in consultation with the director of graduate programs and the department head. Several factors will influence your decision, including laboratory rotations, TA experience, and approval of the graduate advisory committee. A five-member thesis committee is established, typically during the second semester in residence, which includes at least three members of the department faculty and at least one member from outside of the department. The thesis committee will meet with the student at least once per year, usually following their presentation in Graduate Seminar.

Preliminary Exam

Students are encouraged to schedule their comprehensive exam once they have completed the majority of their formal coursework. The exam occurs during the fourth or fifth semester of the program. The preliminary exams are a written component (an original research proposal written by the student) and an oral defense of the proposal. After successful completion, the student is admitted into candidacy.

Thesis Research

After completion of the thesis research, and with the approval of the thesis committee, the student writes a thesis, presents a departmental seminar based on the research, and defends the thesis before the thesis committee. The defense generally occurs at the end of the fifth year in residence.

Graduate School Forms

Students can complete the Ph.D. program within five years. It is essential to set goals for your progress and be aware of the deadlines you must meet and forms you to complete to stay in good standing and graduate. The Graduate School has numerous resources to assist you.

/images/cornell/logo35pt_cornell_white.svg" alt="phd biochemistry courses"> Cornell University --> Graduate School

Biochemistry ph.d. (ithaca), field of study.

Biochemistry, Molecular and Cell Biology

Program Description

The graduate program in the Field of Biochemistry, Molecular and Cell Biology (BMCB) offers a Ph.D. degree only. The goal of our program is to introduce and educate students about the fascination of this scientific area, as well as provide them with the tools necessary to succeed in it. The program offers the opportunity to undertake forefront research with access to the latest technology and equipment. Our program includes both core and flexible coursework that intends to provide a depth and breadth of knowledge in the scientific areas encompassed by the field, develop written communication skills, and inform on research ethics. In addition to coursework, students are required to complete three research rotations in the first year and a one-semester teaching experience (in the second year).

Students in the Field of BMCB can select the Biochemistry PhD or Molecular and Cell Biology PhD degree during the application process. Core curricular requirements are maintained across the BMCB and GGD degree programs. The Biochemistry PhD offers coursework and research opportunities that focus on understanding biochemical mechanisms. More details can be found on the BMCB program website . Degree selection during the application process should be informed by an applicant’s current area of research interest. The flexibility of our training program allows students to move between the BMCB and GGD degree programs at the end of the first year if their research interests evolve.

Contact Information

107 Biotechnology Building Cornell University Ithaca, NY  14853

Concentrations by Subject

• biochemistry

Visit the Graduate School's Tuition Rates page.

Application Requirements and Deadlines

Fall, Dec. 1; no spring admission

Requirements Summary:

Candidates come from many academic backgrounds; however, matriculating students will have commonly majored in one of the many areas of biology, chemistry, or physics. There are no set standards; rather we are searching for applicants who we believe can flourish in the program. The Admissions Committee considers the potential for each applicant to succeed, and past or ongoing research experience is considered an important indicator for success in the program. A holistic review takes into account not only the applicant's academic background and research experience but also the applicant’s accomplishments, character, and life experiences. The academic record (GPA) and research experience are considered alongside the statement of purpose and letters of recommendation. There is not a grade point average minimum or cutoff. We value diversity in all its forms.

• All Graduate School Requirements , including  English Language Proficiency  for all applicants
• Applicants are required to supply the following items:
• Current transcripts, all colleges attended 
• Three letters of recommendation
• Statements of Purpose (Academic and Personal)
• Proof of English language competency
• Curriculum Vitae

Learning Outcomes

Upon completion of a Ph.D., students will be able to:

• Make an original and substantial contribution to the field.
• Demonstrate in-depth knowledge of one area of expertise.
• Demonstrate a broad knowledge of theory and research across several sub-disciplines in the field.
• Learn and follow ethical guidelines for working in the field.
• Write and speak effectively to professional and lay audiences about issues in the field.

Narrow Your Search

Return to Field of Study listing

Clear Filters

Table of Contents

Skip to Content

• Undergraduate Program

Graduate Program

• Be a Biochemist

Other ways to search:

• Events Calendar

Ph.D. Program Information

The major goals of the Ph.D. program in biochemistry are to master known principles and techniques, develop the intellectual tools for collecting and analyzing data, to produce new fundamental knowledge through research, and to be able to effectively communicate the results of original research through oral and written means. 

The academic and other requirements for obtaining a Ph.D. degree are outlined below. Students are responsible for understanding their degree program and fulfilling these requirements. In addition, other pertinent information is provided for course registration, salary support, health insurance, exams, and other topics.

This page is provided as a reference, however, the most complete and up-to-date information for current students can be found in the  Departmental Rules for Advanced Degrees , available in the Helpful Links & Useful Documents section below.

Ph.D. Program Timeline

Arriving on campus.

1 st  year students begin at the University of Colorado Boulder in August with a series of orientations. The department will reach out to students with more detailed instructions prior to the beginning of the semester.

Fall and Spring Semesters of Year 1

The 1 st  year typically consists of laboratory rotations, coursework, and a teaching assistantship.

Laboratory Rotations  – Students participate in four rotations during their first academic year (two during each semester). The rotation lasts ~8-weeks. A rotation is a time when a student previews joining a laboratory by participating in research and attending laboratory events/meetings. The goal of laboratory rotations is to help students find a match with prospective thesis advisor. Biochemistry Ph.D. students can rotate with investigators from across the University but must perform their first rotation with a Biochemistry faculty member.

Coursework  – Students take a number of graduate-level courses within their first year. The most important are two semesters of Biochemistry Core, a class only for Biochemistry Ph.D. students that provides a foundation in principles of biochemistry and essentials for research.

Teaching Assistantship  – Students are typically funded as Teaching Assistants for their 1st year. In subsequent years, students are typically funded by their thesis advisors lab as Research Assistants.

Joining a Lab

1 st  year students typically “join a lab” at the end of Spring Semester of their first year. Joining a lab consists of selecting a thesis advisor (also called mentor) with whom to perform their dissertation research. Labs may begin financially supporting students at this time. Students are encouraged to apply to one of the NIH training grants at the University of Colorado Boulder during this time.

After joining a lab, 2 nd  year students in Biochemistry labs usually no longer teach. This year typically consists of applying for fellowships, taking qualifying exams (also known as COMPs), completing elective coursework, and dissertation research.

Applying for Fellowships  – Biochemistry Ph.D. students are encouraged, but not required, to apply to fellowships to support their stipend and research. Our students have received the National Science Foundation Graduate Research Fellowship (NSF GRFP), National Institutes of Health (NIH) F31 Predoctoral Fellowship, and many other prestigious fellowships.

Qualifying Exams (COMPs)  – During the second year, students take qualifying exams. Passing these exams advances students to Ph.D. Candidacy.

Coursework  – Students complete the remaining coursework required to fulfill their degree. Typically, second year students take electives on specialty topics they choose.

Dissertation Research  – After joining a laboratory, students spend virtually all of their time working with their thesis advisor to perform research for their dissertation.

After advancing to Ph.D. candidacy, students spend virtually all of their time working with their thesis advisor to perform research for their dissertation. Students in Biochemistry convene a Thesis Committee of 3–5 faculty members who meet annually to provide advice and guide students through their degree.

Graduation and Degree Conferral

Students who have fulfilled the degree requirements for a Ph.D. in Biochemistry prepare a written Dissertation/Thesis and hold a Thesis Defense. The average time to completion is 5.5 years.

Ph.D. Degree Requirements

• General Requirements
• Academic & Registration Requirements

All students in the Biochemistry Ph.D. program must be admitted through the University of Colorado Boulder Graduate School. For more information on applying to Biochemistry, see Prospective Graduate Students .

Ph.D. students are required to complete a minimum of 30 hours of approved coursework and 30 dissertation credits.

Students must be enrolled as a full-time, degree seeking graduate students each semester, maintain satisfactory academic performance, and maintain progress toward their degree.

• Graduate Program Overview
• Prospective Graduate Students
• Information, Curriculum, and Requirements
• Meet Our Ph.D. Students
• Interdisciplinary Training Programs
• P2P Mentoring Program

Helpful Links & Useful Documents

Biochemistry resources.

• Annual Committee Meeting Report 
• Biochemistry Handbook Year-by-Year
• Departmental Rules for Advanced Degrees
• Example Written Exams (COMPs)

Graduate School Resources

• Graduation requirements
• Establishing CO residency
• Graduate school rules 
• Doctoral final examination form

Other Resources

• Campus resources for graduate students
• Comprehensive list of crisis resources
• Assistance for students in distress

Biochemistry PhD Program

The application for fall 2025 is now open.

NIH T32 Sponsored Programs

• Training Grant in Cellular, Biochemical & Molecular Sciences
• Graduate Women in Science (GWIS)

The Biochemistry & Molecular Biology (BMB) Program is designed for students interested in obtaining a Ph.D. in Biochemistry. As such, the program represents a group of faculty that may mentor a student pursuing thesis research toward the Biochemistry Ph.D. Students interested in a Ph.D. in Biochemistry should apply to the Biochemistry & Molecular Biology Program for admission to graduate studies at the University of Rochester Medical Center.

Why Choose Our Program?

We offer in-depth coursework and diverse research opportunities that focus on understanding the biochemical mechanisms of life’s critical molecular processes.

World-class research in our laboratories exposes our students to a variety of the latest methods for sophisticated biochemical analysis, including mass spectrometry, crystallography, microcalorimetry, surface plasmon resonance, microarrays, fluorescence activated cell sorting, light scattering, and spectroscopic methods (including fluorescence lifetime and energy transfer measurements), as well as modern methods for cell culture, protein purification, genetic analysis, and reconstitution of biochemical complexes and reactions.

The flexibility of our training program allows students to train in a number of exciting research areas, and often allows students to develop highly effective interdisciplinary collaborations, resulting in cutting edge thesis projects.

What Sets Us Apart

Information on core courses, electives, lab rotations to guide you through our program

We support and guide trainees in several areas of research. Browse research by current program trainees

Mentor Relationships

Meet the faculty mentors and student researchers on our program and see photos of our department family

Discover the Biochemistry PhD Program

All Student Tours

What Our Students Say...

"I chose the University of Rochester graduate program because as an UR undergrad, I had early exposure to the amazing research opportunities and investigators. I knew, for a fact, that I would receive high quality training and experience, making me a competitive candidate in the future. What I enjoy the most about the program is the collaboration and support the BMB program provide for its students. They really go above and beyond to ensure that our needs are met."

Leon Harvey

Upcoming events.

DBB Seminar: “Tackling Functional Elements in the Putative Noncoding Genome: Polyadenylation and Noncanonical Translation”

Zhe Ji, PhD Assistant Professor, Department of Pharmacology, Feinberg School of Medicine; Department of Biomedical Engineering, McCormick School of Engineering, Robert H. Lurie Comprehensive Cancer Center, Northwestern University, Chicago, IL

  Wed, Sep 25 @ 2:00 PM   MC | K-307

All Events »

Latest News

September 5, 2023 Biochemistry Ph.D Student Madeline Jensen Receives Perfect Score on NIH Fellowship Application

June 9, 2023 UR Center for RNA Biology Members and Alumni Attend RNA 2023 in Singapore

May 19, 2023 A Search Engine for mRNA: Algorithm Identifies Optimal Sequences to Improve COVID Vaccines

Connect With Us On Social

Study at Cambridge

About the university, research at cambridge.

• Undergraduate courses
• Events and open days
• Fees and finance
• Postgraduate courses
• How to apply
• Postgraduate events
• Fees and funding
• International students
• Continuing education
• Executive and professional education
• Courses in education
• How the University and Colleges work
• Term dates and calendars
• Visiting the University
• Annual reports
• Equality and diversity
• A global university
• Public engagement
• Give to Cambridge
• For Cambridge students
• For our researchers
• Business and enterprise
• Colleges & departments
• Email & phone search
• Museums & collections
• Course Directory

PhD in Biochemistry

Postgraduate Study

• Why Cambridge overview
• Chat with our students
• Cambridge explained overview
• The supervision system
• Student life overview
• In and around Cambridge
• Leisure activities
• Student union
• Music awards
• Student support overview
• Mental health and wellbeing
• Disabled students
• Language tuition
• Skills training
• Support for refugees
• Courses overview
• Department directory
• Qualification types
• Funded studentships
• Part-time study
• Research degrees
• Visiting students
• Finance overview
• Fees overview
• What is my fee status?
• Part-time fees
• Application fee
• Living costs
• Funding overview
• Applying for University funding
• Doctoral training programmes
• External funding and loans
• Colleges overview
• College listing overview
• Accommodation
• Applying overview
• Before you apply
• Entry requirements
• Application deadlines
• How do I apply? overview
• Application fee overview
• Application fee waiver
• Life Science courses
• Terms and conditions
• Continuing students
• Disabled applicants
• Supporting documents overview
• Academic documents
• Finance documents
• Evidence of competence in English
• AI and postgraduate applications
• Terms and Conditions
• Applicant portal and self-service
• After you apply overview
• Confirmation of admission
• Student registry
• Previous criminal convictions
• Deferring an application
• Updating your personal details
• Appeals and Complaints
• Widening participation
• Postgraduate admissions fraud
• International overview
• Immigration overview
• ATAS overview
• Applying for an ATAS certificate
• Current Cambridge students
• International qualifications
• Competence in English overview
• What tests are accepted?
• International events
• International student views overview
• Akhila’s story
• Alex’s story
• Huijie’s story
• Kelsey’s story
• Nilesh’s story
• Get in touch!
• Events overview
• Upcoming events
• Postgraduate Open Days overview
• Discover Cambridge: Master’s and PhD Study webinars
• Virtual tour
• Research Internships
• How we use participant data
• Postgraduate Newsletter

Primary tabs

• Overview (active tab)
• Requirements
• How To Apply

Students are based in a research group and undertake a research project agreed with the principal investigator (supervisor).  Each student also has additional mentorship support from an advisor and a Postgraduate Thesis Panel.  Although no formal, examined coursework exists, the student's progress is assessed at various stages.  The most important is the First-Year Assessment, which comprises submitting a thesis report and an oral examination assessed by two examiners.  Registration for a PhD is only possible following a satisfactory outcome from the First-Year Assessment.  In their second year, students present their research work in poster form.  In the third year, students present their work in a talk given to the whole Department.

The PhD in Biochemistry is designed to train students to be effective research scientists.  In addition to developing deep knowledge of a subject area and a broader understanding of the relevant field, students will develop skills in:

• identifying experimentally answerable scientific questions and assessing the value of these questions to both science and society
• designing and executing experiments that test these scientific questions
• analysing and interpreting experimental data
• presenting the aims and outcomes of scientific research in both written and oral form
• sourcing, assessing, and critically evaluating scientific literature
• time, data, and resource management
• collaborative and interpersonal professional behaviour

The students will acquire many of these skills within the research group.  Other skills will be developed outside their research group: at the beginning of their first year, students attend an in-house training course of lectures and workshops, and can subsequently continue their training in many areas, choosing from the wide range of courses available within the Postgraduate School of Life Sciences and the University.  Students will also belong to a Peer Research Group, composed of other postgraduate students, within which workshops, student research projects, and scientific literature are presented and discussed.

The University hosts and attends fairs and events throughout the year, in the UK and across the world. We also offer online events to help you explore your options:

Discover Cambridge: Master’s and PhD study webinars - these Spring events provide practical information about applying for postgraduate study.

Postgraduate Virtual Open Days - taking place in November each year, the Open Days focus on subject and course information.

For more information about upcoming events  visit our events pages .

Key Information

3-4 years full-time, 4-7 years part-time, study mode : research, doctor of philosophy, department of biochemistry, course - related enquiries, application - related enquiries, course on department website, dates and deadlines:.

Some courses can close early. See the Deadlines page for guidance on when to apply.

Easter 2025

Michaelmas 2025, easter 2026, funding deadlines.

These deadlines apply to applications for courses starting in Michaelmas 2025, Lent 2026 and Easter 2026.

Postgraduate Admissions Office

• Admissions statistics
• Start an application
• Applicant Self-Service

At a glance

• Bringing a family
• Current Postgraduates
• Cambridge Students' Union (SU)

University Policy and Guidelines

Privacy Policy

Information compliance

Equality and Diversity

Terms of Study

About this site

About our website

Privacy policy

© 2024 University of Cambridge

• Contact the University
• Accessibility
• Freedom of information
• Privacy policy and cookies
• Statement on Modern Slavery
• University A-Z
• Undergraduate
• Postgraduate
• Research news
• About research at Cambridge
• Spotlight on...

• Campus Maps
• Campus Tours
• People Directory
• New Students
• Current Students
• Faculty and Staff
• Brightspace
• Get help with your login
• Faculty & Staff

Explore Programs

Find your program here.

Explore the diverse array of undergraduate, graduate, and professional programs supporting over 200 degrees in 13 faculties at Dalhousie University.

View the glossary for help with language on this page.

Already decided on a program? Learn how to apply .

This website is best viewed in a modern browser with Javascript enabled. While it should still be accessible to older browsers or non-javascript enabled browsers, some functionality may be limited.

www.astate.edu

• Undergraduate
• International
• Early College Programs
• Financial Aid
• University Housing
• Records and Registration

As soon as you set foot on our campus you will know you’ve found the place you can call home.

• Degrees Offered
• Colleges & Departments
• Honors College
• Graduate School
• Online Programs
• Campus Querétaro
• Degree Centers
• Military Science
• NYIT College of Osteopathic Medicine at A-State
• VC for Academic Affairs & Research

Our students are engaged in a wide range of academic pursuits that include degree programs in 160 undergraduate and graduate fields delivered by 6 different colleges.

Visit the Office of Academic Affairs & Research >>

• Student Union
• Leadership Center
• Multicultural Center
• International Programs
• Campus Recreation
• Dining Services
• Student Health Center

A-State offers unlimited possibilities for students to customize their experience while on campus.

Visit Student Affairs >>

• Research & Technology Transfer
• Centers & Institutes
• Sponsored Programs Accounting

Research engages intellectual curiosity, satisfies the thirst for discovery, and provides an outlet for creativity.

• History & Heritage
• Accreditations
• About Jonesboro
• Visit Campus
• Institutional Research & Planning
• Chancellor's Office
• Consumer Information

Marking its first hundred years, Arkansas State University continues to expand in exciting ways.

• Future Students
• Current Students
• Faculty & Staff
• Maps & Directions
• University Police
• Faculty / Staff Directory
• A-State Home »
• Academics »

Biochemistry

College: Beck College of Science and Mathematics

Description

The Bachelor of Science in Biochemistry is where biology and chemistry meet. You'll delve into areas such as structural biology, genetics, and metabolism. Biochemistry is your gateway to understanding the molecular basis of health and disease. With a degree in Biochemistry, you'll be prepared for a wide range of careers, from the medical industry to forensics.

Curriculum and degree requirements can be found in the most current Undergraduate Bulletin.

Visit the bulletins page >>

Request Information Phone: 870-972-3086 Fax: 870-972-3089 Room: 213 Lab Science East Postal Address: PO Box 419 State University, AR  72467

• Open Records
• Appropriate Use
• IT Security
• Accessibility
• Fiscal Transparency

Call Support +91 9614910910

Live chat +91 7076600600, mail support [email protected], search anything.., perm state medical university.

Ulitsa Kuybysheva, 39, Perm, Perm Krai, Russia, 614000

Fees Structure

Eligibility, type of college private, established 1916, affiliated ministry of education of russian federation, mci approved nmc & who, courses offered mbbs.

Perm State Medical University is one of the oldest and most prominent medical universities in Russia. Set up in 1916, the medical university has shown excellent performance in terms of medical education. This renowned medical university is located in a well-connected part of the city making it more accessible for students.

Perm State Medical University is recognized by the WHO & NMC. The medical university has garnered respect among Indian students and other international pupils. This WHO-recognized medical university offers MBBS courses taught by industry experts. The medical graduates from Perm State Medical University have been successfully placed in different medical sectors worldwide.

Any applicant interested in applying to Perm State Medical University for MBBS in Russia have to follow the following admission procedure:

Contact Pice Educare’s admission department.

•  Next, our academic counselors will fill out the application form from Perm State Medical University •  After that, necessary documents need to be furnished by the candidate for the smooth process of admission letter. •  We will then go ahead with the verification process. •  Also, Pice Educare will review the student’s eligibility for successful admission to Perm State Medical University. •  Next, an admission letter from Perm State Medical University will be dispatched for the student. •  Later, the visa and immigration process will begin. •  Now, the students are required to pay the minimum charges for their class. •  Once the verification process comes to an end, the candidate will receive the green signal to start their classes. •  Next, the candidates have to pay the admission charges for the first year. •  Finally, the students will receive flight tickets to chase their dreams.  

Perm State Medical University’s objective is to produce proficient healthcare professionals equipped with sound medical knowledge and a sense of integrity and empathy. With advanced facilities, expert faculties, and a commitment to developed medical research, the medical university offers a rigorous academic environment for the all-round growth of the students.

As Per Academic Session 2023-24

Before being admitted to this famous medical university, one needs to know the fee structure of this university. This will help them to understand the financial responsibility.

[The fee structure provided on this website is for general reference only and may not reflect the exact or current fees. Variations may occur due to factors like college policies and seat allocation. For accurate and up-to-date fee details, please contact our academic counselors directly.]

Medical graduates from Perm State Medical University receive quality education and practical exposure so that they can excel in the future as medical healers. However, candidates must be aware of the career opportunities after getting an MBBS degree from Perm State Medical University. Internship: Perm State Medical University provides a yearlong internship program for future doctors. This not only the students practical exposure to the real world but also gives them compassion toward humanity.

Specialization Degree: After having an MBBS degree, medical healers can pursue higher study to become specialized doctors. This would widen their career opportunity.

General Physician: MBBS graduates can practice in hospitals after completing their MBBS degree from Perm State Medical University. Real-life challenges will help them to sharpen their medical knowledge and place them in a good position in the future.

• Modern Laboratory: Perm State Medical University has research laboratories equipped with modern infrastructure that aims to provide superior practical training for the students. The medical college offers separate research laboratories for Anatomy, Biochemistry, and Physiology.
• Lecture Halls: Perm State Medical University offers four centrally air-conditioned lecture halls with a seating capacity of 180 students at a time. The lecture halls have audio-visual facilities for interactive learning. The medical classes are recorded so that students can attend important lectures from the hostel in case of illness or any other issues.
• Central Library at Perm State Medical University : Perm State Medical University has a chief library and 17 departmental libraries dedicated to medical books, journals, WHO publications, CD ROMs, and Internet services. These are designed in a way that serves the needs of students, and faculties and for research purposes.
• Sports and Games: Perm State Medical University has outdoor and indoor sports facilities for the candidates. They organize outdoor games like Cricket, Football, Volleyball, Basketball, and a 400-meter track for athletics and indoor games like carom, chess, and others. JIMSH has a dedicated sports area for girls equipped with sports like badminton, football, throwball, and basketball.
• Cafeteria: The cafeteria at Perm State Medical University provides delicious and hygienic foods at a reasonable cost. The menus are decorated in a way to gives the essence of home-cooked food and serves the delicacies of Bengal.
• Hostel Facility: Perm State Medical University provides residential areas with modern facilities for girls and boys both. The hostels provide recreational activities too.

Demo Room: Every medical department of Perm State Medical University has its demo room with a seating capacity of 75 students at a time. These demo rooms come with audio-visual facilities and CCTV surveillance.

Students must meet the following eligibility conditions to apply for MBBS in Russia at Perm State Medical University:

●    NEET Qualification is Mandatory ●    The students must have scored at least 50% in Physics, Chemistry, and Biology ●    The student must be 17 years old.

The Perm State Medical University is located in Perm Krai Russia.

The course duration for MBBS at Perm State Medical University is 6 years.

● NEET Qualification is Mandatory ● The students must have scored at least 50% in Physics, Chemistry, and Biology ● The student must be 17 years old.

Yes, it is mandatory

Yes, it is recognized by WHO.

Student Review

Medical college in russia.

Mari State University

Yoshkar-Ola, Mari El Republic, Russia

Orenburg State Medical University

Orenburg Oblast, Russia

Pskov State University

Pskov Oblast, Russia

Omsk State Medical University

Omsk Oblast, Russia

Request for Callback enquire?

Our admission help desk, share this place, admission open for 2024-25 session, how pice educare helps you for your mbbs, md/ms/mds admission journey.

Documentation Process

Medical College Details

Passport & Visa Process

Counselling as per Marks & Budget

Scholarships & Education Loans

Comparison of Medical Colleges

Application & Counselling Process

Seat Booking in Top Private Medical Colleges in India

Seat Booking in Top Medical Colleges in Abroad Countries

Welcome to Pice Educare. In order to provide a more relevant experience for you, we use cookies to enable some website functionality. Cookies help us see which articles most interest you; allow you to easily share articles on social media; permit us to deliver content, jobs and ads tailored to your interests and locations; and provide many other site benefits. For more information, please review our Cookies Policy

• Perm Krai Tourism
• Perm Krai Hotels
• Perm Krai Bed and Breakfast
• Flights to Perm Krai
• Perm Krai Restaurants
• Things to Do in Perm Krai
• Perm Krai Travel Forum
• Perm Krai Photos
• Perm Krai Map
• All Perm Krai Hotels
• Perm Krai Hotel Deals

Trains bypassing Perm-2 - Perm Krai Forum

• Europe    
• Russia    
• Volga District    
• Perm Krai    

Perm Krai Hotels and Places to Stay

• GreenLeaders

Perm State Medical University Russia 2024-25: Fees, Ranking, Admission, Courses, Eligibility etc.

Perm State Medical University Russia is one of Russia’s oldest and largest universities. It was established in 1916 by the government under the Ministry of Education, it offers a 6-year MBBS program in English. The university is recognized by the National Medical Commission of India, the World Health Organization, and other leading medical organizations globally.

Since 1992, Perm State Medical University Russia has had an international student exchange program through the IFMSA. Over 100 specialists have been trained to work successfully in foreign medical institutions in the Middle East, Africa and Asia. More than 3,400 students study at Perm State Medical University Russia every year. 365 clinical interns are trained in 22 specialties, 264 clinical residents are trained in 40 specialties and 94 graduate students are trained in 20 specialties.

[Page Index]

College summary.

Before we complete the college summary, let us look at the essential details of Perm State Medical University Russia .

Wants to Study MBBS Abroad from a top Country with low tution Fees? Subscribe Now!

Affiliation and Recognition

These are the renowned bodies that’ve given a reputation to the Perm State Medical University Russia.

• NMC (National Medical Commission of India).
• WHO (World Health Organization).
• Ministry of Health of the Russian Federation.
• World Directory of Medical Schools (WDOMS).
• Education Commission for Foreign Medical Graduates (ECFMG).
• Registered with the Foundation for Advancement of International Medical Education and Research ( FAIMER ).

The faculties at Perm State Medical University Russia differ from one department to another, encompassing various disciplines and specialities.

• Faculty of Medicine
• Faculty of pediatrics
• Faculty of Dentistry
• Faculty of Medico-prophylactic
• Faculty of Clinical Psychology and Higher Nursing Education
• Faculty of Continuing Professional Education
• Faculty of Pre-University Education and Targeted Education

Courses Offered

Perm State Medical University Russia Courses offer quality medical programs under highly qualified faculty and state-of-the-art infrastructure. It is famous for its undergraduate medical programs. If you want to know about more details the MBBS course check it out below.

Why Study MBBS at Perm State Medical University Russia?

• Recognized by WHO and MCI, allowing graduates to practice medicine internationally.
• Offers a comprehensive MBBS program with experienced faculty and English-medium courses.
• Tuition and living expenses are significantly lower compared to many Western countries.
• Provides hands-on training in well-equipped hospitals and clinics.
• Offers robust student support services and a multicultural learning environment.

Admission Procedure

If you want to take Perm State Medical University admission in 2024, you must qualify for the National Eligibility Entrance Exam (NEET) for Indian students.

Eligibility Criteria

In this section, all the students check the eligibility criteria of Perm State Medical University Russia.

Graphical Representation of Eligibility Criteria

Documents Required

Before admission to Perm State Medical University Russia, please do not forget to carry all these related documents.

• Passport (Minimum 18 months validity).
• 10th Certificate & Mark sheet.
• 12th Certificate & Mark sheet.
• Birth Certificate.
• 10 passport-size Photographs.
• Official Invitation letter from the PSMU.
• Authorization of all documents from the Ministry of External Affairs, New Delhi.
• Legalization of all documents from the Russian Embassy.
• Bank receipt of 1st Year Perm State Medical University fees (required for some universities).
• HIV test documents.

Fee Structure 2024-25

In this section, all the Indian students get information about the Perm State Medical University MBBS fees. Check all the relevant queries regarding fees following this page:  Low fees for MBBS Colleges in Russia .

Ranking 2024-25

According to Edurank, the Perm State Medical University ranking in Russia and Perm State Medical University world Ranking:

Advantages of MBBS in Russia

About Perm City

• Perm is a Russian city located on the banks of the Kama River near the Ural Mountains. It is the administrative centre of Perm Krai, and the fifteenth-largest city in Russia, with a population of over one million residents.
• The city is also a cultural centre and is home to several museums, theatres and universities.
• Perm is also known for its beautiful natural scenery. The city is surrounded by forests and hills, and there are some parks and gardens within the city limits.
• The city has a well-developed transportation infrastructure, including a network of roads, railways and an airport.
• Perm International Airport connects the city to various domestic and international destinations.

Temperature

Contact Details

Perm State Medical University Russia (PSMU) Address: 614000, Perm, st. Petropavlovskaya, 26 Russia

Frequently Asked Questions (FAQs)

What is the duration of the mbbs course at psmu russia.

The MBBS course duration at PSMU is 6 years.

How many Indian students are studying at PSMU Russia?

Around 800 students are studying at Perm Medical University.

Is Perm State Medical University an English medium?

English is the medium of instruction for all international students. All the tests, exams & classes are conducted in English by teachers/ professors/ doctors for Indians.

• Admission Updates,
• Top Collages

You may like to read

• Chuvash State Medical University 2024-25: Courses, Admission, Eligibility, Fees and Ranking etc.
• Ulyanovsk State University Russia 2024-25: Courses, Admission, Eligibility, Fees, Ranking etc.
• Fergana Medical Institute of Public Health 2024-25: Admission, Fees, Ranking, Eligibility etc.
• Samarkand State Medical University Uzbekistan 2024-25: Admission, Fees, Courses, Ranking, Eligibility etc.

About Harsh

Hello, I'm Harsh Begwani, with a year of expertise in MBBS and Ayush courses. I have detailed knowledge of various colleges' fee structures, cutoffs, and intake procedures. If you're looking for insights or assistance in pursuing MBBS or BAMS courses, feel free to comment below—I'm here to help!

Comments (2)

Hello Manish, Looking for MBBS admission in Perm Medical Institute Neet score is 285 Another neet last year score is around 200 How can get admission for both. Can we get at same location Perm college?

You can contact our MBBS Counselling expert at 8859637232 regarding your query.

Leave a Comment Cancel reply

Notify me via e-mail if anyone answers my comment.

Abroad MBBS Update 2024 : Admission Dates, Top College, Fees, Location, Scholarship etc.

Get admission to Top Overseas Universities with Affordable Fees.